Population dynamics, disturbance, and pattern evolution: Identifying the fundamental scales of organization in a model ecosystem

Authored by Thorsten Wiegand, KA Moloney, SJ Milton

Date Published: 1998

DOI: 10.1086/286172

Sponsors: UFZ-Centre for Environmental Research

Platforms: No platforms listed

Model Documentation: Other Narrative Flow charts Mathematical description

Model Code URLs: Model code not found

Abstract

We used auto- and cross-correlation analysis and Ripley's K-function analysis to analyze spatiotemporal pattern evolution in a spatially explicit simulation model of a semiarid shrubland (Karoo, South Africa) and to determine the impact of small-scale disturbances on system dynamics. Without disturbance, local dynamics were driven by a pattern of cyclic succession, where ``colonizer{''} and ``successor{''} species alternately replaced each other. This results in a strong pattern of negative correlation in the temporal distribution of colonizer and successor species. As disturbance rates were increased, the relationship shifted from being negatively correlated in time to being positively correlated-the dynamics became decoupled from the ecologically driven cyclic succession and were increasingly influenced by abiotic factors (e.g., rainfall events). Further analysis of the spatial relationships among colonizer and successor species showed that, without disturbance, periods of attraction and repulsion between colonizer and successor species alternate cyclically at intermediate spatial scales. This was due to the spatial ``memory{''} embedded in the system through the process of cyclic succession. With the addition of disturbance, this pattern breaks down, although there is some indication of increasing ecological organization at broader spatial scales. We suggest that many of the insights that can be gained through spatially explicit models will only be obtained through a direct analysis of the spatial patterns produced.
Tags
spatial pattern South-africa Species-diversity Alpine abies forests Arid karoo Seedling establishment Nitrogen availability Serpentine grassland Gopher disturbance Germination sites